A white lion of the line of the Bister Kings, powerful lions with commanding voices. Apparently their voices have the ability to make any animal obey them and they can communicate with every animal. Because Madara is white he was ostracized from his family and wasn't allowed to rule as a Bister King. When every human was suddenly turned into an animal he tried to rule the world.
In the end, he sees the error of his ways and decides that coexisting is the best solution. Also, Madara was revealed to be a female lion.[/spolier]